Regillus in 496 BC, were defeated by the Veientes in the Battle of the Cremera in 477 BC, the Sabines in an unnamed battle in 449 BC, the Aequi in the Battle of Mount Algidus in 458 BC, the Aequi and Volsci in 446 BC, in the Battle of Corbio, in 446 BC the Aurunci in the Battle of Aricia, the Capture of Fidenae in 435 BC and the Siege of Veii in 396 BC, and the Capture of Antium in 377 BC. After defeating the Veientes, the Romans had effectively completed the conquest of their immediate Etruscan neighbours, as well as secured their position against the immediate threat posed by the tribespeople of the Apennine hills. In the meantime, it also affected the agriculture as well as diet regime of the empire. Since the enlargement the population in Apennine peninsula had increased and led to certain changes in agriculture, such as switch to goat breeding from cattle, indicating higher levels of protein supply in the diet which played a crucial role in stature of the locals. However, Rome still controlled only a very limited area and the affairs of Rome were minor even to those in Italy and Rome's affairs were only just coming to the attention of the Greeks, the dominant cultural force at the time. At this point the bulk of Italy remained in the hands of Latin, Sabine, Samnite and other peoples in the central part of Italy, Greek colonies to the south, and the Celtic people, including the Gauls, to the north. Celtic invasion of Italia (390–387 BC) By 390 BC, several Gallic tribes had begun invading Italy from the north as their culture expanded throughout Europe. Most of this was unknown to the Romans at this time, who still had purely local security concerns, but the Romans were alerted when a particularly warlike tribe, the Senones, invaded the Etruscan province of Siena from the north and attacked the town of Clusium, not far from Rome's sphere of influence. The Clusians, overwhelmed by the size of the enemy in numbers and ferocity, called on Rome for help. Perhaps unintentionally the Romans found themselves not just in conflict with the Senones, but their primary target. The Romans met them in pitched battle at the Battle of the Allia around 390–387 BC. The Gauls, under their chieftain Brennus, defeated the Roman army of around 15,000 troops and