discovered at Border Cave in KwaZulu-Natal in 2012. Historical evidence shows that certain San communities have always lived in the desert regions of the Kalahari; however, eventually nearly all other San communities in southern Africa were forced into this region. The Kalahari San remained in poverty where their richer neighbours denied them rights to the land. Before long, in both Botswana and Namibia, they found their territory drastically reduced. Genetics Various Y chromosome studies show that the San carry some of the most divergent (oldest) human Y-chromosome haplogroups. These haplogroups are specific sub-groups of haplogroups A and B, the two earliest branches on the human Y-chromosome tree. Mitochondrial DNA studies also provide evidence that the San carry high frequencies of the earliest haplogroup branches in the human mitochondrial DNA tree. This DNA is inherited only from one's mother. The most divergent (oldest) mitochondrial haplogroup, L0d, has been identified at its highest frequencies in the southern African San groups. In a study published in March 2011, Brenna Henn and colleagues found that the ǂKhomani San, as well as the Sandawe and Hadza peoples of Tanzania, were the most genetically diverse of any living humans studied. This high degree of genetic diversity hints at the origin of anatomically modern humans. A 2008 study suggested that the San may have been isolated from other original ancestral groups for as much as 50,000 to 100,000 years and later rejoined, re-integrating into the rest of the human gene pool. A DNA study of fully sequenced genomes, published in September 2016, showed that the ancestors of today's San hunter-gatherers began to diverge from other human populations in Africa about 200,000 years ago and were fully isolated by 100,000 years ago. Ancestral land conflict in Botswana Much aboriginal people's land in Botswana, including land occupied by the San people (or Basarwa), was conquered during colonisation. Loss of land and access to natural resources continued after Botswana's independence. The San have been particularly affected by encroachment by majority peoples and non-indigenous farmers onto their traditional land. Government policies from the 1970s transferred a significant area of traditionally San land to majority agro-pastoralist tribes and white settlers Much of the government's policy regarding land tended to favor the dominant Tswana peoples over the minority San and Bakgalagadi. Loss of land is a major contributor to the problems facing Botswana's indigenous people, including especially the San's eviction from the Central