for the Taira clan, a call for arms was sent up by Minamoto no Yorimasa and Prince Mochihito. Not knowing who was behind this rally, Kiyomori called for the arrest of Mochihito, who sought protection at the temple of Mii-dera. The Mii-dera monks were unable to ensure him sufficient protection, so he was forced to move along. He was then chased by Taira forces to the Byōdō-in, just outside Kyoto. The war began thus, with a dramatic encounter on and around the bridge over the River Uji. This battle ended in Yorimasa's ritual suicide inside the Byōdō-in and Mochihito's capture and execution shortly afterwards. It was at this point that Minamoto no Yoritomo took over leadership of the Minamoto clan and began traveling the country seeking to rendezvous with allies. Leaving Izu Province and heading for the Hakone Pass, he was defeated by the Taira in the battle of Ishibashiyama. However he successfully made it to the provinces of Kai and Kōzuke, where the Takeda and other friendly families helped repel the Taira army. Meanwhile, Kiyomori, seeking vengeance against the Mii-dera monks and others, besieged Nara and burnt much of the city to the ground. Fighting continued the following year, 1181. Minamoto no Yukiie was defeated by a force led by Taira no Shigehira at the Battle of Sunomatagawa. However, the "Taira could not follow up their victory." Taira no Kiyomori died from illness in the spring of 1181, and around the same time Japan began to suffer from a famine which was to last through the following year. The Taira moved to attack Minamoto no Yoshinaka, a cousin of Yoritomo who had raised forces in the north, but were unsuccessful. For nearly two years, the war ceased, only to resume in the spring of 1183. Turning of the tide In 1183, the Taira loss at the Battle of Kurikara was so severe that they found themselves, several months later, under siege in Kyoto, with Yoshinaka approaching the city from the north and Yukiie from the east. Both Minamoto leaders had seen little or no opposition in marching to the capital and now forced the Taira to flee the city. Taira no Munemori, head of the clan since his father Kiyomori's death, led his army, along with the young Emperor Antoku and the Imperial regalia, to the west. The cloistered emperor Go-Shirakawa defected to Yoshinaka. Go-Shirakawa then issued a mandate