A B The lipid structures characterized above are similar to what has been found previously, and are broadly in line with the literature consensus despite the greater complexity of lipid composition. A key question is whether the membrane dynamics are similarly replicated across different lipid types. From Fig. , it is clear that the mean squared displacement is larger for cis lipids than for trans lipids, and so we should expect the higher diffusion for the disordered all-cis membranes. Lipid lateral diffusion is a readily accessible metric that can be compared to both experiment and varying simulation models, with a wide range of characterized values depending on the membrane. Some report self diffusion coefficients from simulation of around 0.06×10 -8 cm 2 , 62 while others report substantially higher diffusion coefficients of 3-8×10 -8 cm 2 /s, although the membrane size in simulated bilayers can dramatically change the measured lipid lateral diffusion. In our simulations, we observe diffusion on the higher end of this range (Fig. ), in line with current expectations for a membrane in the L d phase.