Skeletal muscle properties are determined by a functionally cooperative set of genes specific to the spatiotemporal requirements of each muscle . In the process of muscle development, growth, and maturation, genes are expressed through regulation at the transcriptional, post-transcriptional, and translational levels, resulting in metabolite generation by multistep enzymatic reactions . Even after tissue growth and maturation, the content of muscle metabolites is altered through gene expression and regulation based on physiological requirements, as observed in muscle type specification . Skeletal muscle metabolite composition is a major factor in characterizing muscle properties, and thereby the nutritional and sensory properties of the meat; thus, metabolite composition provides important phenotypic information .