In the equation: 𝑛 𝑖 is the amount of lipid type i, 𝑁 𝐺 is the number of lipid types in the system, C is the environmental concentration of lipid i, 𝑘 𝑒𝑛𝑡𝑟𝑦,𝑖 and 𝑘 𝑒𝑥𝑖𝑡,𝑖 are the basal entry and exit rates of lipid type i into and from a homogeneous cluster (100% lipid i), 𝑓 𝑗 is the fraction of the composition that pertains to modulator lipid type j, and 𝛽 𝑒𝑛𝑡𝑟𝑦,𝑖𝑗 and 𝛽 𝑒𝑥𝑖𝑡,𝑖𝑗 are the rate modification parameters of modulator j over probe i, taken from the derived β-matrices (Figures and).