Members of the millipede order Platydesmida have intriguing qualities such as egg brooding, and other quasisocial traits, and are almost exclusively associated with various fungal taxa, which some consume in unusual circular social aggregations called pinwheels . All of the species of the Platydesmida with characterized chemical secretions, and its evolutionary sister Polyzoniida , have disparate heterocyclic terpenoid alkaloids, including Brachycybe lecontii from the eastern U.S. that produces 1 and Gosodesmus claremontus from California and Oregon that produces 3 ) (Fig. ). Interestingly, despite B. lecontii and Brachycybe petasata both being known from the eastern U.S., even cooccurring on the same decaying logs at some sites, B. petasata is not the closest evolutionary sister of B. lecontii.