Combining multiple engineered traits in a single plant is becoming increasingly feasible, allowing for more complex and comprehensive crop improvements. This approach offers several advantages and is rapidly advancing the field of plant biotechnology. Synergistic trait combinations can lead to greater improvements than individual modifications alone, as different traits can complement and enhance each other's effects . For instance, stacking genes for drought tolerance with those for improved nutrient use efficiency can result in crops that perform better under a wider range of environmental conditions. Multigene pathway engineering has enabled the introduction or modification of entire biosynthetic pathways for complex metabolites or agronomic traits . This has been particularly valuable in biofortification efforts and in engineering plants to produce highvalue compounds.