Due to the existence of large thermal fluctuations in biological systems and in light-harvesting complexes , we proceed to determine the energy ladder within the FCP complex as average over an ensemble of conformations. To this end, we have performed a 1 ns-long QM/MM MD simulation. Then, excitation energy calculations were carried out for the resulting conformations to determine the energy fluctuations along this QM/MM MD trajectory. The DFTB3 method with the 3OB-f parameter set was utilized for the ground state simulations, whereas the TD-LC-DFTB scheme with the OB2 parameter set was employed for the excita- These findings strongly suggest that the excitation energy within this FCP complex might be funneled towards the Chl-c2 pigment, i.e., an energy transfer from the Chl-c2 pigment towards the Chl-a pool is, on average, highly unlikely. In recent experimental studies, pumpprobe and two-dimensional spectroscopy has revealed an ultra-fast energy from the Chl-c2 pigments towards the Chl-a pool . These experiments were made, however, for the diatom C. meneghiniana with a different (structurally so far unresolved) FCP complex than the one probed herein. Thus, no direct comparison can be made, but a re-analysis of these experimental findings might be in order.