The values that are within the estimated experimental error range of ±0.02 39 are coloured white. Statistical error in simulations is not considered here because it is approximately order of magnitude smaller than the experimental error. Order parameters of hydrogens attached to the same carbon were averaged, except for the C2 carbon of the oleate chain (whose order parameters forked) for which dierences for both the larger and smaller values were calculated, and the average of these dierences is shown in the C2 column. The dependence of the area per phospholipid (APL) on CHOL concentration follows the trends in thickness inversely in general (bottom right panel of Fig. ), yet provides curious dierences between force elds at the physiologically relevant CHOL concentration range.