Solids storage and analyses for protein-to-N ratio and carbohydrate content were performed as in Bradley et al. 2021. Briefly, protein content was estimated by multiplying the elemental nitrogen content by a conversion factor that represents the ratio of N content to protein. Conversion factors were determined by analysis of amino acid residuals; amino acid profiling was performed by Bio-Synthesis, Inc. (Lewisville, Texas, USA). Total monomeric carbohydrate content of lyophilized solids was determined after a two-step acid hydrolysis of the complete biomass. The hydrolysate was neutralized, filtered, and the monosaccharide concentration was quantified against glucose standards. Solids crude lipid content was quantified as in Gardner-Dale et al. Briefly, crude lipids from lyophilized solids were extracted using an adaptation of the Folch method and a 2:1 (v/v) chloroform:methanol solvent mixture. After the extraction, sodium chloride solution was added to bring the final mixture to a 8:4:3 ratio of chloroform:methanol:sodium chloride. The mixture was centrifuged, resulting in a biphasic system; the bottom phase containing the crude lipids was transferred to weighting dishes to be measured gravimetrically after the carrier solvent evaporated.