<prompt> Bhala ukuqhubeka kwalesi sigaba - Wazalelwa futhi wafundiswa eBrno, eCzechoslovakia, futhi wasinda ekuQothulweni Kwesizwe ngokufakwa ku-Kindertransport, futhi wachitha iminyaka yempi eNgilandi. Wabuyela eCzechoslovakia ngemuva kwempi, wathola iziqu zobudokotela futhi waqeqeshwa njengodokotela wezingane lapho. Isiqu sakhe sobudokotela eNyuvesi yaseBrno sasithi Genetika isoamylas: Studie nového lidského polymorfismu. ngesiNgisi: I-Genetics of Isoamylases: Study of the New Human Polymorphism ngo-1967. Ngemva kokuhlasela iCzechoslovakia  <response>: ngo-Agasti 1968, waphunyuka okwesibili waya eNgilandi, lapho asebenza khona noLionel Penrose eKennedy-Galton Centre for Medical and Community Genetics eLondon ngokulimala kwengqondo. Waqokwa njengelungu lezikole e-University of Wisconsin-Madison ngo-1975, lapho asebenza khona esikhungweni sayo sokucwaninga ngokukhubazeka kokukhula komuntu, i-Waisman Center, ekuxilongweni kwangaphambi kokubeletha nasekululekeni kwezakhi zofuzo. Waba nguprofesa ohlonishwayo ngo-2003. 