<prompt> Qedela umusho olandelayo: Wazalelwa futhi wafundiswa eBrno, eCzechoslovakia, futhi wasinda ekuQothulweni Kwesizwe ngokufakwa kuKindertransport, futhi wachitha iminyaka yempi eNgilandi. Wabuyela eCzechoslovakia ngemuva kwempi, wathola iziqu zobudokotela futhi waqeqeshwa njengodokotela wezingane lapho. Isiqu sakhe sobudokotela eNyuvesi yaseBrno sasithi Genetika isoamylas: Studie nového lidského polymorfismu. ngesiNgisi: I-Genetics of Isoamylases: Study of the New Human Polymorphism ngo-1967. Ngemva  <response>: ukuhlasela kweCzechoslovakia ngo-Agasti 1968, wabaleka okwesibili waya eNgilandi, lapho asebenza khona noLionel Penrose eKennedy-Galton Centre for Medical and Community Genetics eLondon ngokulimala kwengqondo. Waqokwa njengelungu lezikole e-University of Wisconsin-Madison ngo-1975, lapho asebenza khona esikhungweni sayo sokucwaninga ngokukhubazeka kokukhula komuntu, i-Waisman Center, ekuxilongweni kwangaphambi kokubeletha nasekululekeni kwezakhi zofuzo. Waba nguprofesa ohlonishwayo ngo-2003. 