 Ramba uchinyora ndima iyi - Litopterna kubva patsoka yakatsetseka iboka rakatsakatika remhuka dzinoyamwisa dzine mahwanda dzakawanikwa munguva yeCenozoic. Urongwa uhwu ndehumwe hwezvikamu zvishanu zvikuru zvezvipembenene zvekuSouth America izvo zvakapa kondinendi iyi mhuka yakasiyana nedzimwe kusvika paGreat American Biotic Interchange. Kufanana nedzimwe mhuka dzinoyamwisa dzomuSouth America, ukama hwadzo nemamwe mapoka emhuka dzinoyamwisa hwakanga husina kujeka kwenguva refu, asi uchapupu hwemajini uye hweproteomic hunoratidza kuti hama dzadzo dzepedyo dziri kurarama ndiPerissodactyls nguruve dzisina zvigunwe kusanganisira mabhiza, mhembwe, uye tapirs, uye kuti litopterns yakabatana zvikuru notoungulates, rimwe boka rakapararira reSouth America ungulates. 