 Ramba uchiverenga ndima ino - South American native ungulates, inowanzo kupfupikiswa seSANU, imhuka dzakafa dzakafanana nemapapiro emhuka dzine hukama hunopokana dzaive dzekuSouth America pamberi peGreat American Biotic Interchange. Dzinosanganisira mapoka makuru mashanu anowanzorongwa semapokaAstrapotheria, Litopterna, Notoungulata, Pyrotheria, uye Xenungulatapamwe nemamwe mapoka, akadai seDidolodontidae neKollpaniinae. Zvave zvichirongwa kuti dzimwe kana kuti nhengo dzose dzeboka iri dzinoumba boka, rinonzi Meridiungulata, kunyange zvazvo ukama hwezvipembenene zveSouth America huchiri kusagadziriswa. Mapoka makuru maviri ezvipembenene zvomuSouth America, notoungulates uye litopterns, ndiwo mapoka bedzi akaramba aripo pashure pepakati peMiocene. Mhatsa shomanene bedzi dzemarotoungulates nelitopterns ndidzo dzakapukunyuka kutozosvikira pakutsakatika kwePleistocene yokupedzisira. 