 Chikepe chacho chakagadziridzwazve muNorfolk, Virginia kubva musi wa24 Ndira kusvika musi wa7 Kurume 1942, apo chakagamuchira radar yezvombo zvepfuti yeType 273 uye mamwe maOerlikon matanhatu. Pashure pokupedza kugadziridza kwake, Devonshire akatumirwa kunobatana ne4th Cruiser Squadron yeEastern Fleet uye akaendesa mutungamiri kubva kuCharleston, South Carolina, kuenda kuFreetown, Sierra Leone munzira. Musi wa25 Kubvumbi, ngarava iyi yakafambisa ngarava kubva kuDurban, South Africa kuenda kuMadagascar sechikamu cheOperation Ironclad, iyo yakatangwa kuti isapinda muchitsuwa ichi. VaFrance veVichy vakanga vabvumira mauto eJapan kushandisa nzvimbo dzendege muFrench Indochina kutanga kurwisa kwemhepo uko kwakanyudza ngarava huru dzeRepulse nePrince of Wales, naizvozvo zvaibvira zvikuru kuti vaizobvumira maJapan kupinda muzvivako zvemauto paMadagascar kudai vakanga vachikumbira. Mukupera kwa1942, Devonshires single four-inch AA pfuti dzakatsiviwa nemaputi maviri-pfuti emidziyo yeMark XVI pfuti yeiyo caliber imwechete. Ngarava yacho yakaramba iri kuFar East kusvikira muna May 1943, ichifukidza ngarava dzehondo kubva kuSuez kuenda kuAustralia. Panguva ino, zvombo zvake zvina zvemachina.50-caliber uye matanhatu e-Oerlikon akaiswa panzvimbo yeimwe miviri ye-octuple-pounder maviri uye gumi nemaviri-Oerlikon. Akadzokera kumusha mwedzi iwoyo ndokutanga kugadziridza kwenguva refu kwakapfuurira kusvikira 20 March 1944. Munguva yokugadziridza uku, X turret uye imwe twin Oerlikon mount yakatsiviwa nemamwe maviri e-octuple maviri-pounder mount uye masere e-Oerlikon mount. Radar suite yake yakagadziridzwa neType 281B ichitsiva Type 281 uye huwandu hwakazara hwean anti-aircraft artillery radars hwakaiswa. Kuti agadzirise midziyo yose mitsva, catapult yake yakabviswa. 